Sequences record the spread of pathogens
images by Trevor Bedford
Human seasonal influenza viruses
slide by Trevor Bedford
Influenza viruses evolve to avoid human immunity
Vaccines need frequent updates
Enterovirus D68 -- with Robert Dyrdak, Emma Hodcroft & Jan Albert
Non-polio enterovirus
Almost everybody has antibodies against EV-D68
Large outbreak in 2014 with severe neurological symptoms in young children (acute flaccid myelitis)
Another outbreak in 2016
Outbreaks tend to start in late summer/fall
Several reports of EV-D68 outbreaks last fall (201 AFM cases in the US in 2018)
How does EV-D68 spread? Does it change?
Whole genome deep sequencing of Enterovirus D68
Amplified in 4 overlapping segments
Illumina sequenced to high coverage
Dyrdak et al, biorxiv
iSNV frequency accuracy and sequencing errors
iSNV frequencies reproducible above 1%
background at around 1/1000
Dyrdak et al, biorxiv
Infections with multiple variants
A set of iSNVs at very similar frequencies in full linkage
Suggest infection with two related variants
3 out of 50 samples: Implies high prevalence
Dyrdak et al, biorxiv
Carbapenemase producing bacteria
Reserve antibiotics used to treat MDR bacteria
Introduced in the 1980ies
Resistance spread rapidly
Resistance is mediated by several distinct beta-lactamases
→ pressing public health problem
→ fascinating instance of genes sweeping the globe by horizontal transfer
Tracking bacteria by sequencing
Illumina → millions of short reads (<500bp)
Too short to bridge repetitive elements
→ assemblies are fragmented into 100s of "contigs"
Problem: all the important bits are flanked by repetitive/mobile elements
(really terrible example)
Images: illumina.com, github.com/rrwick
Long-read sequencing of Carbapenemase producing bacteria
Contigs with drug resistance genes ~1-6 genes → no phylogenetic resolution
long-read assemblies give full length plasmids
tracking via synteny and structural diversity, not SNPs
→ we need to reconstruct spread from genome structure evolution
Noll et al, biorxiv, 2018
Synteny alignments of Carbapenemase containing loci
Structural changes resolve evolutionary relationships
Different KPC alleles are found on the same backgrouond
Identical KPC alleles are found on different backgrounds
Similar plasmids are spread across MLSTs and species boundaries
Noll et al, biorxiv, 2018
Summary
Timely data sharing + automated analysis allows near real-time tracking of influenza
Such analyses provide important input for vaccine strain selection
Sequencing, analysis, and dissemination can be rapidly set-up for emerging pathogens
Bacterial pathogens come with a special set of challenges
Fascinating instance of evolution beyond SNPs
Acknowledgments -- nextstrain
Trevor Bedford
Colin Megill
Pavel Sagulenko
Sidney Bell
James Hadfield
Wei Ding
Emma Hodcroft
Sanda Dejanic
John Huddleston
Barney Potter
Acknowledgments -- Enterovirus
Robert Dyrdak
Jan Albert
Lina Thebo
Emma Hodcroft
Bert Niesters (Groningen)
Randy Poelman (Groningen)
Elke Wollants (Leuven)
Acknowledgments -- Bacteria
Wei Ding
Nicholas Noll
Eric Ulrich
Adrian Egli (at USB)
With-in host diversity
Above 0.5%, iSNVs are biological
Most samples have few iSNVs, three had more than 20
Dyrdak et al, biorxiv